In the wake of Virginia being shot, the crew must regroup to face their off-kiltered new adversaries, Mac and Melba Lovestone, so they can run Bayside Rapture Casino like the bosses they are.

Desna and her crew start working at the casino and discover the Lovestones are involved in some illegal business with a very important person.

After discovering Mac and Melba’s surprise gift in the salon, Desna and the crew go to their “fundraiser” to retrieve information to use against them.

After Desna discovers Mac and Melba have done something horrible to a loved one, she realizes just how far she is willing to go to save him.

Desna decides her trust issues are rooted in being abandoned by her father. Her world is turned upside down after seeking him out. The Lovestones retaliate against Desna for getting in the way.

After successfully leveraging what she knows about Governor Patel to win her brother back, a victorious Desna returns to the casino, only to learn that Dean is more entranced under Mac and Melba's spell than she realized.

Desna works to separate Dean from Mac and Melba. Quiet Ann gets the crew deeper into trouble.

The crew makes elaborate plans to pay back the casino's debts only to find out they have a long way to go before they are out of the woods.

Desna and the crew have to tie up loose ends for the Triads if they want to stay alive. Meanwhile, guilt pushes Toby to do the unexpected, putting the entire crew in jeopardy.

Just when the crew thought that they could boss up, fate takes a turn on Desna and Roller. As the walls close in on Desna's Queendom, she is forced to make a choice no one would have expected.

Desna and the girls struggle to keep hope alive while still working under the thumb of the new Russian regime, spearheaded by Riva. Desna remains in the dark about her gangster/gyno boyfriend, Ruval. Virginia tells Dean that she's pregnant.

Desna, now pinned under the stiletto-tipped nail of Riva's sister, Zlata, must come up with an idea to generate more customers for the clinic. Quiet Ann helps Dean and Virginia get to a very important appointment.

Desna's relationship with Ruval deepens when he introduces her to his mother, Matilde. Polly grows concerned when Marnie continues to pull away from her. Jenn, still depressed by Bryce's recent indiscretion, falls off the wagon.

Desna, in an attempt to cozy up to Matilde, Ruval's mother, puts together a spa day for her and a couple of young girls from his charity. Quiet Ann gets a surprise visit from her brother, who convinces her to attend a family dinner.

Desna continues to grow closer to Zlata, despite the crew's increasing concern as her relationship with Dr. Ruval intensifies. Meanwhile, Dean gets an opportunity to showcase his talents and Uncle Daddy gets his mojo back.

Faced with a dilemma, Desna and Dean take a road trip to seek guidance from a childhood mentor. Back in Palmetto, Jenn must lean on Polly and Quiet Ann for help with a personal crisis. Virginia struggles to run the salon on her own.

When Desna with the help of Uncle Daddy and her crew plots revenge on those who betrayed her, she realizes that vengeance comes at a high cost.

Desna is offered a deal with the Russian devil, aka Zlata, that's more intriguing than nail art on acrylics. Polly and Quiet Ann get dealt some hard-ass betrayal from their baes while Jenn's affair comes back to haunt her.

Polly gets an unexpected visit from her long-lost sister that sends her spinning out of control. Uncle Daddy and Roller join forces with their Dixie Mafia brethren and make a plan to take out the Russian and Haitian mafias for good.

As a hurricane approaches, Desna proceeds with the plans for her big day, keen for the the opportunity to settle all scores. Opposing forces converge at a lavish golf resort, determined to prevail, but will Desna and crew survive the storm?

Desna and her girls take pride in doing nails at the Nail Artisans of Manatee County, but amidst painting hooves and dreaming of a better life, she finds herself entangled in a money laundering scheme with the Dixie Mafia.

Following their entanglement with Roller, Desna and Virginia desperately prepare a cover story to tell the girls and the Dixie Mafia. Desna's dream salon is falling out of her grasp.

As Virginia's lies deepen, Desna plays catch-up covering their tracks. Desna still has a small shot at her dream salon, and with the help of her crew, she is determined to get it.

Haunted by Roller, Desna is desperate to get out from under Uncle Daddy's thumb. Bryce drifts deep into the Dixie Mafia. Polly reunites with a former gold-digging "friend," while Quiet Ann is in a lady-mess.

Desna and Jennifer may be Dr. Ken's only hope, as Bryce wrongly accuses him of a heinous crime. Virginia and Desna are forced to go to Polly for help to rid the world of two horrible birds from Desna's past, with one well-aimed stone.

All of our girls go to their new fancy salon for the first time. They aren't met with the kindness of strangers. Desna and crew decide to compete in Florida's premiere nail design competition for a big cash prize.

After the big nail competition, Desna may finally get out from Uncle Daddy's grasp. Jennifer struggles knowing Desna's dark secret, while Polly discovers her previously unknown maternal instincts.

With a shocking revelation affecting the entire crew, Jennifer finds unexpected solace in her square-dancing instructor. Polly uses all of her powers of bad for good, for the first time, while Quiet Ann is torn about a big decision.

Desna goes searching for the truth with her crew right behind her, save for Jennifer, who is learning how to cope as a Dixie Mafia wife. When Desna's sleuthing hits pay dirt, she goes to the only person who can save her. Time is running out.

Following a harrowing escape from Roller's grasp, Desna is on the run. Roller is close behind, running from the Russians. In the end, only Desna can save Jennifer, Roller, and her crew with a sacrifice greater than any she's ever had to make.