Guests: Glenn Close and John Waters. Henry Louis Gates, Jr. discovers the privileged lineages of Glenn Close and John Waters.

Guests: Andy Cohen and Nina Totenberg. Henry Louis Gates, Jr. introduces Andy Cohen and Nina Totenberg to their survivor ancestors.

Henry Louis Gates, Jr. explores the roots of actor Jane Lynch and comedian Jim Gaffigan, revealing the Irish American experience through their families.

Guests: Tony Shalhoub and Christopher Meloni. Henry Louis Gates, Jr. reveals the immigrant roots of actors Tony Shalhoub and Christopher Meloni, introducing ancestors who came to America to build a better life.

Guests: Pharrell Williams and Kasi Lemmons. Henry Louis Gates, Jr. helps musician Pharrell Williams and filmmaker Kasi Lemmons uncover extraordinarily rare first-person accounts of their enslaved ancestors.

Guests: Clint Black and Rosanne Cash. Henry Louis Gates, Jr. uncovers the remarkably diverse backgrounds of country music icons Clint Black and Rosanne Cash.

Guests: John Lithgow and Maria Hinojosa. Henry Louis Gates, Jr. traces ancestors of actor John Lithgow and journalist Maria Hinojosa who thrived in North America long before the birth of the United States.

Guests: Gretchen Carlson and Don Lemon. Henry Louis Gates, Jr. examines how journalists Gretchen Carlson and Don Lemon were able to overcome biases in their careers, drawing parallels to relatives who met profound challenges of their own.

Guests: Audra MacDonald and Mandy Patinkin. Henry Louis Gates, Jr. investigates the family histories of Broadway stars Audra MacDonald and Mandy Patinkin, discovering ancestors whose struggles laid the groundwork for their success.

Guests: Lewis Black and Roy Wood, Jr. Henry Louis Gates, Jr. provides comedians Lewis Black and Roy Wood, Jr. with accounts of hardship in their family trees which may have led them to each find humor in adversity.

Actresses Isabella Rossellini, Anjelica Huston and Mia Farrow explore their family histories, generations beyond their cinematic forebears.

Actors Melissa McCarthy and Eric Stonestreet retrace their ancestral journeys.

Sterling K. Brown, Jon Batiste and Sasheer Zamata learn about the unexpected places their ancestors called home.

The unexpected family trees of Queen Latifah and Jeffrey Wright challenge preconceptions about America's past.

Exploring the Jewish heritages of actor Jeff Goldblum, radio host Terry Gross and comic Marc Maron.

Sigourney Weaver, Justina Machado and Amy Ryan explore their ancestry.

Tracing the family histories of scientists Harold Varmus, Francis Collins and Shirley Ann Jackson.

Questlove, S. Epatha Merkerson and Ava DuVernay explore the unexpected places where their ancestors were scattered by slavery.

Marisa Tomei, Jimmy Kimmel and John Turturro learn about the sacrifices their ancestors made to bring their families from Italy to America.

The family histories of Laura Linney, Lisa Ling and Soledad O'Brien reveal tricksters, scoundrels and criminals.

Guests: Diane von Furstenberg, Narciso Rodriguez, RuPaul. Henry Louis Gates, Jr. steps into the world of fashion and meets three icons of style — Diane Von Furstenberg, Narciso Rodriguez, and RuPaul Charles — introducing ancestors who were just as audacious as they are.

Guests: Téa Leoni, Joe Madison. Henry Louis Gates, Jr. uses DNA detective work to solve mysteries in the family trees of actor Téa Leoni, and radio host Joe Madison, introducing each of them to parents and grandparents whose names they’ve never heard before. (Téa Leoni originally appeared in S04E08. Joe Madison originally appeared in S05E10.)

Guests: Kehinde Wiley, Julianne Moore, Bill Hader. Henry Louis Gates, Jr. explores the roots of actor Julianne Moore, comedian Bill Hader, and painter Kehinde Wiley, revealing how their ancestors’ military service left an indelible mark both on their families and on their country. (S03E06 was also titled "War Stories." Kehinde Wiley originally appeared in S05E09. Julianne Moore originally appeared in S03E06. Bill Hader originally appeared in S03E04.)

Guests: Lupita Nyong’o, Lidia Bastianich, Scarlett Johansson. Henry Louis Gates, Jr. meets actors Scarlett Johansson and Lupita Nyong’o, and chef Lidia Bastianich — three women whose families crisscrossed the globe to escape oppression and find opportunity, leaving them with questions about their relatives who stayed behind. (Lupita Nyong’o originally appeared in S04E07. Lidia Bastianich originally appeared is S03E09. Scarlett Johansson originally appeared in S04E05.)

Guests: Gayle King, Jordan Peele, Issa Rae. Henry Louis Gates, Jr. explores the family secrets of journalist Gayle King, film director Jordan Peele, and comedian Issa Rae, introducing them to ancestors who raise profound questions about the shape and meaning of their family trees.

Guests: Nancy Pelosi, Norah O'Donnell, Zac Posen. Henry Louis Gates, Jr. helps Congresswoman Nancy Pelosi, news anchor Norah O’Donnell and fashion designer Zac Posen explore their immigrant roots, retracing the journeys of their ancestors who arrived in the United States with little more than a dream--and revealing why that dream is still so inspiring today.

Henry Louis Gates, Jr. helps actor Andy Samberg and author George R. R. Martin answer some family mysteries when DNA detective work uncovers new branches of their family trees. The DNA analysis upends family history and reveals new relatives.

Henry Louis Gates, Jr. helps actors Felicity Huffman and Michael K. Williams discover a wealth of hidden family history. Both grew up knowing little about their biological fathers, and seeing their lost ancestry restored proves deeply empowering.

Henry Louis Gates, Jr. shows journalists Christiane Amanpour, Ann Curry and Lisa Ling that the stories within their own family trees are every bit as compelling as the news stories they have been covering for the world.

Henry Louis Gates, Jr. explores with Marisa Tomei, Sheryl Sandberg and Kal Penn the tremendous challenges faced by their immigrant forebears. From Italy, Russia and India to America, their histories show success could take generations to achieve.

Henry Louis Gates, Jr. delves deep into the roots of two African-American guests, actor S. Epatha Merkerson and athlete and television personality Michael Strahan. Both discover unexpected stories that challenge assumptions about black history.

Henry Louis Gates, Jr. explores the diverse family histories of politicians Paul Ryan, Tulsi Gabbard and Marco Rubio. They each see a shared vision of the great American melting pot reflected in their ancestors’ stories.

Henry Louis Gates, Jr. shows comedians Seth Meyers, Tig Notaro and Sarah Silverman that their family trees are filled with people whose struggles laid the groundwork for their success. Gates also reveals to each one news of an unexpected DNA cousin.

Henry Louis Gates, Jr. explores the family stories of filmmaker Michael Moore and actors Laura Linney and Chloë Sevigny—three people whose distant ancestors overcame great hardships in ways that resonate with their lives today.

Henry Louis Gates, Jr. shares the family histories of director Alejandro G. Iñárritu, iconoclastic performance artist Marina Abramović and painter Kehinde Wiley. These visionary artists find their identities challenged—and affirmed.

The family stories of actor Ty Burrell, and radio host Joe Madison. Also a look at how a summer camp for children is using genetic genealogy to help young people better understand their burgeoning identities.

Join Larry David and Bernie Sanders, who learn they have more in common than they thought.

Guests: Carly Simon, Christopher Walken, and more. In this episode, three guests each learn about a grandparent whose real identity and background had been a mystery to them. Along the way, two of them also discover that their close relatives were on the wrong side of history.

Guests: Ted Danson, Mary Steenburgen and William H. Macy. Three mainstays of modern-day Hollywood discover family legacies that predate the United States itself.

Guests: Ta-Nehisi Coates, Ava DuVernay and Janet Mock. Three guests who have helped to redefine Black America in the last decade find their identities challenged as they learn about their family origins.

Guests: Scarlett Johansson, Paul Rudd and John Turturro. In this episode, three guests explore the tremendous challenges faced by their immigrant forebears.

Guests: Bryant Gumbel, Tonya Lewis-Lee and Suzanne Malveaux. Three African-American guests delve deep into their family trees, discovering unexpected stories that challenge our assumptions about black history.

Guests: Lupita Nyong’o, Carmelo Anthony and Ana Navarro. Three guests explore how their family trees were shaped by political turmoil and violence, discovering sometimes unexpected ancestry along the way.

Guests: Gaby Hoffmann and Téa Leoni. Two guests whose lives have been shaped by family mysteries are introduced to biological ancestors they never knew they had thanks to genetic detective work.

Guests: Questlove, Dr. Phil and Charlayne Hunter-Gault. Three guests of disparate backgrounds dig into their Southern roots, where slavery and its aftermath shaped families both black and white.

Guests: Amy Schumer and more. Guests who have found fame mining their family stories for comedy learn about ancestors who overcame immense suffering.

Discover the unsolved mysteries behind the family stories of political organizer Donna Brazile, actor Ty Burrell and artist Kara Walker as they learn how the legacy of slavery has shaped their identities.

Explore the shared Irish ancestry of three powerful social commentators of our times, Bill Maher, Soledad O'Brien, and Bill O'Reilly.

Learn how the ancestors of Maya Rudolph, Shonda Rhimes and Keenen Ivory Wayans struggled for freedom, and how each of these cultural trailblazers gain a new understanding of how they fit into this proud trajectory.

Uncover a history of tragedies and triumphs in the families of three of America’s funniest men–Jimmy Kimmel, Norman Lear and Bill Hader—as they learn how comedy truly has its place in all aspects of life.

Discover how the ancestors of business mogul Richard Branson and architects Maya Lin and Frank Gehry took audacious risks to create opportunities, and how their luck, ingenuity and chutzpah was passed on to these three visionaries.

Patricia Arquette, Senator John McCain, and Julianne Moore explore their genealogical heritage with Henry Louis Gates, Jr.

Uncover family mysteries about two legends of hip hop, Sean Combs and LL Cool J, through the use of DNA technology, which reveals information that shakes their very foundations.

Discover how Neil Patrick Harris, Gloria Steinem and Sandra Cisneros are connected to pioneers who broke new ground and paved the way for their modern day descendants to continue shaping our culture.

Discover how actress Julianna Margulies, author Azar Nafisi and chef Lidia Bastianich are bound together by their ancestors’ singular and deeply human desire to preserve their most cherished traditions.

Learn how Dustin Hoffman and Mia Farrow, who last shared the screen together in 1969, also share a history of tragic deaths in their families that played major roles in shaping future generations.

Three famous guests — Stephen King, Gloria Reuben and Courtney B. Vance — know almost nothing about their fathers’ histories.

Billie Jean King, Derek Jeter and Rebecca Lobo learn about their family histories. King discovers that her grandmother was an orphan; Derek Jeter investigates his ancestors' lives as slaves; and Lobo finds out that an ancestor was forced to flee Spain because of his democratic ideals.

Ken Burns learns that his Southern ancestors include Confederate soldiers and a slave owner; Anderson Cooper learns that an ancestor on his father's side was killed by one of his slaves; and Anna Deavere Smith learns about her great-grandfather, who was a free black man and former conductor on the Underground Railroad.

Actor Ben Affleck discovers his mother's past as a Freedom Rider in 1964, NAACP President Ben Jealous learns how far his great-grandfather would go to keep his family whole, and actress Khandi Alexander learns about her grandfather's death. This episode is not available from pbs because actor Ben Affleck had persuaded Gates to omit information about his slave-owning ancestors. In the investigation, PBS said that producers violated network standards by letting Mr. Affleck have “improper influence” and “by failing to inform PBS or WNET of Mr. Affleck’s efforts to affect program content.”

Chef Tom Colicchio learns the hardships his family endured in Northern Italy; chef Ming Tsai traces his roots back 2,000 years; chef Aarón Sánchez learns his Mexican roots include Spaniards, Africans and Native Americans.

Nas learns about his slave ancestors; Angela Bassett discovers that her family history includes an emancipation story; and Valerie Jarrett, a senior adviser to President Obama, learns about the free people of color in her family's past

Tony Kushner learns how the Holocaust impacted his ancestors; Carole King delves into the origins of her family name; and Alan Dershowitz discovers that an Hassidic synagogue started by his great-grandfather played a secret role in World War II.

Deepak Chopra, Sally Field and Sting’s ancestry demonstrates the extraordinary influence of the British Empire during the 18th and early 19th century.

Tina Fey, George Stephanopoulos and David Sedaris, three quintessential voices in American life, descend from ancestors that were part of the wave of Greek immigrants that flooded America’s shores in the early decades of the 20th century.

Secrets encoded in their genomes challenge these guests’ ideas about their families’ histories and their identities today. This episode features Jessica Alba, Anderson Cooper, Valerie Jarrett, Governor Deval Patrick and more.

Their European immigrant ancestors blazed unconventional trails in America, from capturing British ships for the American Revolution to crossing racial barriers in slave-era Louisiana. Generations later, as children growing up in New Orleans, Harry Connick, Jr. and Branford Marsalis found a deep and abiding friendship through their common love of jazz and of the city itself. In this hour, we trace the turbulent and contradictory history of the city of New Orleans through the family stories of these two fascinating men.

In this episode, we feature two African American politicians from different generations and opposite backgrounds. John Lewis grew up in a sharecropping family in rural Georgia, while Cory Booker was raised in an affluent, all-white New Jersey suburb. Although both men have devoted their lives to the betterment of African-American people, neither of them knows much about their own ancestors. In this episode, we introduce Booker to his white great-grandfather, a man he never knew, and move Lewis to tears over the extraordinary ambitions and accomplishments of his slave ancestors..

What's in a name? Well, a lot, at least when it comes to piecing together your family history. For former slaves, choosing a last name was one of their first acts of freedom. For Jewish immigrants, it was a way to fit in their new country. Whatever the reason for a name change, it can make the process of learning about one's ancestors difficult, if not impossible. In this episode, we unearth missing links in the family histories of media legend Barbara Walters and educational superstar Geoffrey Canada. Walters did not know her father's real last name. Canada did not know the name of his grandfather. Both of them had been unable to access their history... until now.

What do Kevin Bacon and Kyra Sedgwick have in common? They are both famous actors and both descend from prominent American families that have been in this country since its inception. But they share something else, too: both had ancestors who were early opponents of slavery. Bacon's Quaker ancestors repudiated slavery long before the rest of the country, in 1780. And Sedgwick's ancestor Theodore Sedgwick argued the freedom case of Elizabeth Freeman, also known as "Mumbet," in 1781 -- which helped bring an end to slavery in Massachusetts. We reveal this fact - and many others - to Sedgwick and Bacon in this episode, and learn quite a bit about slavery in the North in the process.

Pastor Rick Warren, Rabbi Angela Buchdahl and Sheik Yasir Qadhi are of three different faiths, all with complex family histories that profoundly shaped their religious beliefs. Our research of their roots unearthed a story about the spiritual foundations of this country, an unrelenting struggle to achieve our ideal of religious freedom and tolerance, but also the difficulties sometimes of holding onto one's faith and still feeling like an "authentic" American.

Many American descend from a dense soups of European ethnicities; we could call them mutts, with ancestral roots across every country in Europe. Actors Robert Downey, Jr. and Maggie Gyllenhaal are textbook examples of Americans with a rich tapestry of European immigrant stories making up their family trees. We delve in to their deep American roots in early colonial communities, and we meet their Eastern European Jewish ancestors, who share almost identical migration stories From the Mayflower to Ellis Island, we journey through centuries of immigration that shaped and built our melting pot nation.

Samuel L. Jackson, Condoleezza Rice and Brown University President Ruth Simmons have each climbed to the pinnacle of their profession, yet each started life as a second class citizen in the Jim Crow south. In this episode we will use DNA to investigate family mysteries: where do they come from in Africa, and who are the white men in their family trees?

The three guests in this episode are all children of first- or second-generation immigrants and share the peculiar burdens of that heritage. In an episode that crisscrosses the planet, from India to Korea to Poland, catch a glimpse of three distinct yet oddly overlapping experiences of families leaving their homes and becoming American.

Most African Americans struggle to trace their ancestors beyond Emancipation; slavery erased names and family ties with brutal efficiency. But what about the descendants of the handful of free black people who evaded bondage during that terrible time? Musician John Legend and comedian Wanda Sykes discover the extraordinary stories of the free black ancestors they never knew about, while Professor Gates himself and his 98-year-old friend Margarett Cooper delve into the mysteries shrouding the free people of color in their family trees.

Michelle Rodriguez, Adrian Grenier and Linda Chavez all share Spanish colonial roots, yet each views their own identity very differently. In this episode we ask what it means to be Hispanic-and find that the answer lies in the tangled histories of European, Native American and African peoples. Crisscrossing Mexico, Spain, the Caribbean, and the Southwest, Professor Gates reveals stories of ancestral Conquistadors, Indian rebels, and "Crypto-Jews" (Spanish Jews who converted to Catholicism to survive the Inquisition, yet continued to practice their religion ...